Chapter 6 No.6

Botanical notes connected with the foregoing Journal.

(February 19th.-The finest view of the hills from Upper Assam is obtained on a reach or turn of the river just above Palankar, the river bending to the NNE. Snow is plentifully seen on one back range from the Sugar-loaf peak. Another reach shortly after presents a fine view of the Burrampooter chasm, terminated by the rugged peak so distinctly seen from Suddiyah, due east. This view might be chosen, as a general characteristic of the Scenery of Upper Assam.

It embraces the Mishmee mountains to the left, the higher peaks of which are covered with perpetual snow. These lie to the NNE. of Beesa. To the east, is the continuation of the Himalaya, to the South-east and South, the Patkaye, and Naga ranges; the whole forming a panorama, rarely if any where surpassed in beauty. Temperature. of the river at 6 A.M. 67°

Musa. Many flowers from the axil of a bract; no bractioles interspersed, hence we may expect racemose or spicate partial inflorescences. The perianth is unilateral, 5 cleft, the two smaller segments, which are intermediate, being internal, or belonging to a different series. Within this petaloid perianth is a membranous one, together with a boat-shaped bracteolate body, entire. The stamens are five, evidently opposite to the segments of the petaloid perianth, staminibus adnatis, the sixth is not developed, but is rudimentary, and exceedly minute, opposite to the bracteoid body. The carpella three, alternate as they ought to be with the last series of stamina, and hence they are opposed to the larger and outer segments of the petaloid perianth, but this last point deserves further examination.

The base of the bracteoid sepal is filled with a gelatinous, sweet, transparent, unicoloured ? fluid.

I am unaware whether this explanation has occurred to any body else.

It is curious as compared with Scitamine?, in which the posticous stamen is alone fully developed. Pl. 1. Fig. 3. a. bracteoid body, b. sterile stamen, c.c.c. outer series, d.d. inner ditto.

The fact of the outer smaller lacini? belonging to a second series is not very apparent, but is corroborated by the evidently internal situation of the bracteoid scale, and by the evidently elevated lines visible in the inner.

(April 3rd, 1837.-On march towards the Serpentine mines) the face of the perianth, corresponds to these smaller lacini?.

April 7th.-Thunbergia grandiflora has the pedicels of its flowers twisted, or not twisted, according to the situation of the flowers. Thus if the flower be so situated that the raceme has the direction of the axis, or in other words is erect, the pedicel is straight, but if the raceme, as generally happens, be pendulous, the twisting of the pedicel is resorted to, to secure the flower that situation which it would have, were the raceme erect.

The above is obvious in flowers which from elongation of the axis of inflorescence, have fasciculate or aggregate flowers. An obvious inference is, that the twisting of the pedicel is not of generic, nor of specific importance; and that it is capable of being produced artificially.

This resupination is not uncommon in the order; it is most evident in Thunbergia coccinea, in which the racemes are always pendulous. There is nothing, at least in this species, in the situation of the genitalia to account for the resupination.

Pedicelli demum apicem infra articulati, the inflorescence of this order is always centrifugal, the partial axis being invariably as well indeed as the general, disposed to dichotomy. Hence the very common presence of three bracte? to each flower, the central one presenting the leaf from whose axil the partial branch springs.

Stipul?-if the analogy of these be difficult to ascertain, the structure and functions would appear to be as of leaves, in addition to the function of protection. In most cases they are certainly not double organs; in Naucleace? they are apparently so. Can this be explained by supposing them to form a bud with four scales, the scales instead of being imbricate, being on one plane. Stipell? of Leguminos? are certainly single; these being all probably stipulate plants, are to be considered as having terminal buds, the buds being either totally, or partially protected by the stipul?. The difficult nature of ochre? of Polygone? is certainly to be acknowledged, but they are similar to those of Costus, and hence not stipul?, but an extension of the margin of the vaginate petiole, from which veins are prolonged into it; the functions of these are not stomatose, since they are membranous, the veins being the only green parts.

I see no reason why the stipul? of Ros? are not to be considered as belonging to, or dilatations of the petiole. They have no distinct vascular fascicles to indicate a distinct origin. And further, in Lowea no stipul? exist.

Jonesia: pedicellis apice articulatis, basi bracteolatis, ideoque infloresc. magis composita esse debet; laciniis anticis? coroll?? perianth compositum, binatum pr?bentibus, emarginatio et situs stam 5ti rudiment. Staminis laciniis alternatis? basi in annulum, seriem 2 indicantem coalit. {111} The situation of the stamens is somewhat obscure, the two lowermost however alternate with the segments, the two intermediate being sometimes sub-opposite.

Of course if they be opposed, the perianth will be referrible to a calyx if not to a corolla.

Lepidostachys or Scepa. Fruit dicarpillary, stigmata four, hence they are placentary not costoid. bilocular, loculis dispermis, ovula 2 pend; 1 abortiv. semiunceum, testa vix arillus obsacuit clause lutescens carnosa et ab nuclei inter adh?ren. Rad. sup. embryo junior viridis.

Stipul? cad. Gemmam oblegent.

Homaline?, Calycis; lacini? 4, petal 4, Glandul? 4 totidem sepalis opposit?. Connat; stamin 4, petal opposita; styli 4. Ovar non ext.

Arbor magna. foliis alternis stipulatis, panicul? racemoso-axillares, Flores minut. viridescent. Pet. et sep. fimbriat. ?stiv. imbricat.

Clematis has semina pendula.

The stipul? of Ficus obviously belong not to the leaves, their insertion taking place ? a line above that of the petiole. Hence they belong as obviously to the elongation of the axis above the leaf; their coloration is curious, especially as they are green when young. Their vernation is conduplicate and plicate.

Combretum presents several points in common with Rhamne?; valvate calyx, and tendency to want of petals; to El?agne? in calyx and furfuraceous scales; a decandrous Rhamne? would differ but little in flowers from Combretum.

My idea of the origin of stigmata is proved to be correct by a Phyllanthus, the carpella of which are ovuliferous below, the upper part being fleshy, the stigmata are two to each, obviously corresponding to the placentary inflexions, while the sinus terminating the dorsal suture is totally naked; it is this which should bear the stigma if Lindley's view were correct.

The true place of Moringa seems to be near Xanthophyllum with which genus it has some remarkable points of resemblance, witness the papilionaceous corolla; unilocular stamina, their situation, ovary, placentation, and lastly glandulation.

To this Lindley has made an approximation by placing the order near Violarie?. Its chief difference from Polygale?, is habit, foliation, and the perigynous insertion of corolla and stamina, and consequent union of the sepals. As in Xanthophyllum there is no albumen.

(An additional Xanthop. which until to-day I have always taken for a Leguminosa.)

Tamarindus cal 4 partitus, sepals 2, superiorib. connatis. Pet. 3, vexillo, sepalo postico composit; opposit; stamen tria; sepalis 3, inferior opposita. Stylus ?stivation deflexus.

Pedicelli apice articulat. Folii petiol. basi articulat. Stipul? minim? stipell?.

In Jonesia, there are no petals. Humboldtia comes near Tamarindus, through H. Brunonis, which agrees in calyx and petals.

Thorns of Prionites, what are they? They are axillary, and yet buds are produced between them and the axis. They have no connection with the leaves. Were it not for the buds above alluded to, I should say that they were abortive branches (bearing one pair of leaves) reduced to spines.

Olacine?. Certainly in habit, corolla, etc. Olacine? are allied to Aurantiace?, but they are nearer akin to Santalace?. The processes are indubitably modified stamina, with a great tendency to irregularity; in one species from Tagoung only three fertile, and five sterile stamina were observed: the three fertile generally, but not invariably, alternate with the petals.

To Santalace? they approach in processes, valvate corolla, and placentation, also to Loranthace?.

Eight stamina thus accounted for; when two opposed to petals, belong to outer series-also single one.

In Punica, the structure of the ovaria is highly curious. We find the bottom of the tube is occupied by two cells, partially filled with ovula, which are attached both to the axis and to the base, as well as to the lower part of the outer paries of each cell; so far, it does not depart from the order, for in Aplexus the placentation is tolerably similar.

Above these two, are 4-5 cells, filled with ovula, which are attached entirely to the outer wall of each cell, but the placent? however would seem to have an obvious connexion with the axis, although this is very doubtful.

The formation of the stigma decidedly indicates a binary formation of carpella.

If these 4 upper cells are 4 constantly, and the base of the ovary is as constantly two celled, then the explanation is sufficiently obvious, though different from that given by Lindley. {113}

First, we have in the bottom from which the mere structure of an ovary is deduced, the normal dicarpellary structure, and there is in addition a tendency in excess toward a parietal placentation.

The anomalous formation arises first from parietal placent? being produced to the axis, and from spurious growth from the sides of the ovary also meeting in the axis, by which the ovula are divided into four bundles.

Lindley's view seems to be questionable, because as in all cases the styles and stigmata are more permanent than ovaries, there should be as many styles, etc. as ovaries. 2nd, because according to this view the placental suture of the carpella would be turned from the axis, (look at Pomace?,) although his view of Pomace? being right would indicate an additional affinity with Mespilus, etc. which it does in habit and abbreviated lateral branches.

Are all Myrtace? dicarpellar?

The true nature of the case is pointed out in the instance cited by Lindley of a permanent variety of apple, which has 14 cells and 14 styles! With regard to Nicotiana and Nolana; have these one or two rows of carpella?

TECTONA.

Arbores, trunco crasso, cito ramoso, cortice albido, l?vi, tenui.

Folia siliceo-aspera, inflorescent dichotoma.

Calyx ?stiv. valvat. cor infundibul, subregularis laciniis, 5 rotundatis, demum reflexis ?stivat. laciniis super 2, omnino exterior, facies barbato-villosa.

Anther? longit dehiscent, stylus stigma simplex.

Pubescentia stellata.

Modo Asclepiade?, corolla rotata.

GMELINA VILLOSA.

Lab. super. ?stivat. omnino exterior fl. axi fere paralleli, pedicell apice bibracteolat.

Cal. minim. 5 dentat.

Cor. infundibul campanul. bilab; 4 partit. stigma bilabiat-lab infer longiore.

AROIDEUM.

Radix maxime napiformis, undique radiculas exserens, et superne e centro spadicem. Spadix pedunculum 3-uncial terminans, basi squamis magnis membranaceis, lineari-oblongis stipatus sursum in corpus fungoiden, capitatum, maximum, purpureo-sanguineum, superficie rugose dilatata.

Ovar bilocul, diovulat.

Medio antheras bipoross confertissimas, sessiles, numerosas, basi ovaria distantiora gerens.

Ovaria fusco-purp, stylus elongatus clavatus, stigma clavato, capitat.

Odor-floris pr?serti marcescentis pessimus.

Katha in sylvis aridis.

The fruit of Lagerstramia grandiflora can, I think, be explained by assuming it to consist of several carpella, which by not becoming united near the axis, leave an irregular shaped space in the centre; the placent? are fleshy, the ovule inserted all around. This view does not take into consideration the situation of the stigmata. The deeper sulci visible externally correspond to the inflexions of the carpellary leaves; in addition to this, the centre of the dorsum of each of these is marked with a line. {114})

            
            

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